The cat (Felis catus), also known as the domestic cat or housecat[5] to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests. Cats have been associated with humans for at least 9,500 years,[6] and are currently the most popular pet in the world.[7] Owing to their close association with humans, cats are now found almost everywhere in the world.
Cats are similar in anatomy to the other felids, with strong, flexible bodies, quick reflexes, sharp retractable claws, and teeth adapted to killing small prey. As crepuscular predators, cats use their acute hearing and ability to see in near darkness to locate prey. Not only can cats hear sounds too faint for human ears, they can also hear sounds higher in frequency than humans can perceive. This is because the usual prey of cats (particularly rodents such as mice) make high frequency noises, so the hearing of the cat has evolved to pinpoint these faint high-pitched sounds. Cats also have a much better sense of smell than humans.
Despite being solitary hunters, cats are a social species and use a variety of vocalizations, pheromones and types of body language for communication. These include meowing, purring, trilling, hissing, growling, and grunting.[8]
Cats have a rapid breeding rate. Under controlled breeding, they can be bred and shown as registered pedigree pets, a hobby known as cat fancy. Failure to control the breeding of pet cats by spaying and neutering and the abandonment of former household pets has resulted in large numbers of feral cats worldwide, with a population of up to 60 million of these animals in the United States alone.[9]
As The New York Times wrote in 2007, "Until recently the cat was commonly believed to have been domesticated in ancient Egypt, where it was a cult animal",[10] but a study that year revealed that the lines of descent of all house cats probably run through as few as five self-domesticating African Wildcats (Felis silvestris lybica) c. 8000 BC, in the Near East.[4] The earliest direct evidence of cat domestication is a kitten that was buried alongside a human 9,500 years ago in Cyprus.[11]
Contents
[hide] 1 Nomenclature and etymology
2 Taxonomy and evolution
3 Genetics
4 Anatomy
5 Physiology
6 Senses
7 Health 7.1 Diseases
7.2 Poisoning
8 Behavior 8.1 Sociability
8.2 Grooming
8.3 Fighting
8.4 Hunting and feeding
8.5 Play
8.6 Reproduction
9 Ecology 9.1 Habitats
9.2 Impact on prey species
9.3 Impact on birds
10 Cats and humans 10.1 Domesticated varieties 10.1.1 Coat patterns
10.1.2 Body types
10.2 Effects on human health
10.3 Indoor scratching
10.4 Waste
11 Feral cats
12 History and mythology
13 See also
14 References
15 Bibliography
16 External links
Nomenclature and etymology
The word cat derives from Old English catt, which belongs to a group of related words in European languages, including Welsh cath, Spanish gato, French chat (French pronunciation: [ʃa]),[12] Basque katu, Byzantine Greek κάτια kátia, Old Irish cat, Frisian and Dutch kat, German Katze, Armenian katu, and Old Church Slavonic kotka. The ultimate source of all these terms is Late Latin catus, cattus, catta "domestic cat", as opposed to feles 'European wildcat'. It is unclear whether the Greek or the Latin came first, but, like Arabic qiṭṭ and Nubian kàdíís, they were undoubtedly borrowed from a word in an Afro-Asiatic language akin to Berber kaddîska, meaning 'wildcat'.[13] The term puss (as in pussycat) may come from Dutch poes or from Low German Puuskatte, dialectal Swedish kattepus, or Norwegian pus, pusekatt, all of which primarily denote a woman and, by extension, a female cat.[14]
Classification based on human interaction[15]
Population
Food source
Shelter
Socialized
Pedigree
Fed by owner
Human homes
Yes
Pet
Fed by owner
Human homes
Yes
Semi-feral
General feeding
Buildings
Yes
Feral
General feeding/foraging
Buildings
No
Pseudo-wildcat
Foraging/hunting
None
No
While wildcats are the ancestral species from which domestic cats are descended, there are several intermediate stages between domestic pet and pedigree cats and these entirely wild cats. The semi-feral cat is a cat that is not owned by any one individual, but is generally friendly to people and may be fed by several households. Feral cats are associated with human habitations and may be fed by people or forage in rubbish, but are wary of human interaction. Pseudo-wildcats are descended from domestic cats, but now tend to live entirely independently from people.[15]
A group of cats is referred to as a "clowder", a male cat is called a "tom" (or a "gib", if neutered), and a female is called a "molly" or "queen". The male progenitor of a cat, especially a pedigreed cat, is its "sire", and its female progenitor is its "dam". An immature cat is called a "kitten" (which is also an alternative name for young rats, rabbits, hedgehogs, beavers, squirrels and skunks). In medieval Britain, the word kitten was interchangeable with the word catling.
A cat whose ancestry is formally registered is called a pedigreed cat, purebred cat, or a show cat. In strict terms, a pure-bred cat is one whose ancestry contains only individuals of the same breed. A pedigreed cat is one whose ancestry is recorded, but may have ancestors of different breeds. Cats of unrecorded mixed ancestry are referred to as domestic longhairs and domestic shorthairs or commonly as random-bred, moggies, mongrels, or mutt-cats.
Taxonomy and evolution
Main article: Cat evolution
The wildcat Felis silvestris is a close relative and possible ancestor of the domestic cat.
The Felids are a rapidly evolving family of mammals that share a common ancestor only 1015 million years ago,[16] and include, in addition to the domestic cat, lions, tigers, cougars, and many others. Within this family, domestic cats (Felis catus) are part of the genus Felis, which is a group of small cats containing seven species.[1][17] Members of the genus are found worldwide and include the Jungle Cat (Felis chaus) of southeast Asia, the African Wildcat (Felis silvestris lybica), the Chinese Mountain Cat (Felis bieti) and the Arabian Sand Cat (Felis margarita).[18]
All the cats in this genus share a common ancestor that probably lived around 67 million years ago in Asia.[19] Although the exact relationships within the Felidae are still uncertain,[20][21] both the Chinese Mountain Cat and the African Wildcat are close relations of the domestic cat and are both classed as subspecies of the Wildcat Felis silvestris.[4][20] As domestic cats are little altered from wildcats, they can readily interbreed. This hybridization may pose a danger to the genetic distinctiveness of wildcat populations, particularly in Scotland and Hungary.[22]
The domestic cat was first classified as Felis catus by Carolus Linnaeus in the tenth edition of his Systema Naturae of 1758.[2][23] However, because of modern phylogenetics, domestic cats are now usually regarded as another subspecies of the Wildcat Felis silvestris.[4][23][24] This has resulted in mixed usage of the terms, as the domestic cat can be called by its subspecies name, Felis silvestris catus.[1][4] Wildcats have also been referred to as various subspecies of F. catus,[24] but in 2003 the International Commission on Zoological Nomenclature fixed the name for wildcats as F. silvestris.[25] The most common name in use for the domestic cat remains F. catus, following a convention for domesticated animals of using the earliest (the senior) synonym proposed.[25] Sometimes the domestic cat is called Felis domesticus[26] or Felis domestica,[23] the term coined by German naturalist Johann Christian Polycarp Erxleben in 1777. These are not valid taxonomic names, and Linnaeus' binomial takes precedence.[27]
Cats have either a mutualistic or commensal relationship with humans. However, in comparison to dogs, cats have not undergone major changes during the domestication process, as the form and behavior of the domestic cat are not radically different from those of wildcats, and domestic cats are perfectly capable of surviving in the wild.[28][29] This limited evolution during domestication means that domestic cats tend to interbreed freely with feral cats, which distinguishes them from other domesticated animals.[15] However, several natural behaviors and characteristics of wildcats may have preadapted them for domestication as pets.[29] These traits include their small size, social nature, obvious body language, love of play and relatively high intelligence;[30] they may also have an inborn tendency towards tameness.[29]
There are two main theories about how cats were domesticated. In one, people deliberately tamed cats in a process of artificial selection, as they were useful predators of vermin.[31] However, this has been criticized as implausible, because there may have been little reward for such an effort: cats do not carry out commands and, although they do eat rodents, other species such as ferrets or terriers may be better at controlling these pests.[4] The alternative idea is that cats were simply tolerated by people and gradually diverged from their 'wild' relatives through natural selection, as they adapted to hunting the vermin found around humans in towns and villages.[4]
Genetics
Main article: Cat genetics
Blue-eyed cats with white fur have a high incidence of genetic deafness.[32]
The domesticated cat and its closest wild ancestor are both diploid organisms that possess 38 chromosomes[33] and roughly 20,000 genes.[34] About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors.[35] The high level of similarity among the metabolisms of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats in the study of the human diseases.[36][37]
An example of a mutation that is shared among all felines, including the big cats, is a mutant chemosensor in their taste buds that prevents them from tasting sweetness, which may explain their indifference to fruits, berries, and other sugary foods.[38] In some breeds of cats congenital deafness is very common, with most white cats (but not albinos) being affected, particularly if they also have blue eyes.[32] The genes responsible for this defect are unknown, but the disease is studied in the hope that it may shed light on the causes of hereditary deafness in humans.[39]
Since a large variety of coat patterns exist within the various cat breeds, the cat is an excellent animal to study the coat genetics of hair growth and coloration.[40] Several genes interact to produce cats' hair color and coat patterns. Different combinations of these genes give different phenotypes. For example, the enzyme tyrosinase is needed to produce the dark pigment melanin and Burmese cats have a mutant form that is only active at low temperatures, resulting in color appearing only on the cooler ears, tail and paws.[41] A completely inactive gene for tyrosinase is found in albino cats, which therefore lack all pigment.[42] Hair length is determined by the gene for fibroblast growth factor 5, with inactive copies of this gene causing long hair.[43]
Purebreds normally have amber eyes, while nonpedigrees are more likely to have green eyes
The Cat Genome Project, sponsored by the Laboratory of Genomic Diversity at the U.S. National Cancer Institute Frederick Cancer Research and Development Center in Frederick, Maryland, aims to help the development of the cat as an animal model for human hereditary and infectious diseases, as well as contributing to the understanding of the evolution of mammals.[37] This effort led to the publication in 2007 of an initial draft of the genome of an Abyssinian cat called Cinnamon.[34] The existence of a draft genome has led to the discovery of several cat disease genes,[34] and even allowed the development of cat genetic fingerprinting for use in forensics.[44]
Anatomy
Diagram of the general anatomy of a male
Domestic cats are similar in size to the other members of the genus Felis, typically weighing between 4 kilograms (8 lb 13 oz) and 5 kilograms (11 lb 0 oz).[20] However, some breeds, such as the Maine Coon, can exceed 11 kilograms (25 lb). Conversely, very small cats (less than 1.8 kilograms (3 lb 15 oz)) have been reported.[45] The world record for the largest cat is 21.297 kilograms (46 lb 15.2 oz).[46] The smallest adult cat ever officially recorded weighed around 1.36 kilograms (3 lb).[47] Cats average about 2325 centimeters (910 in) in height and 46 centimeters (18.1 in) in head/body length (males being larger than females), with tails averaging 30 centimeters (11.8 in) in length.[48]
Cats have 7 cervical vertebrae like almost all mammals, 13 thoracic vertebrae (humans have 12), 7 lumbar vertebrae (humans have 5), 3 sacral vertebrae like most mammals (humans have 5 because of their bipedal posture), and a variable number of caudal vertebrae in the tail (humans retain 3 to 5 caudal vertebrae, fused into an internal coccyx).[49] The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.[50] Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones, which allow them to pass their body through any space into which they can fit their heads.[51]
Skull
The cat skull is unusual among mammals in having very large eye sockets and a powerful and specialized jaw.[52] Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death.[53] Compared to other felines, domestic cats have narrowly spaced canine teeth; which is an adaptation to their preferred prey of small rodents, which have small vertebrae.[53] The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively.[54]
Cats, like dogs, are digitigrades. They walk directly on their toes, with the bones of their feet making up the lower part of the visible leg.[55] Cats are capable of walking very precisely, because like all felines they directly register; that is, they place each hind paw (almost) directly in the print of the corresponding forepaw, minimizing noise and visible tracks. This also provides sure footing for their hind paws when they navigate rough terrain. Unlike most mammals, when cats walk, they use a "pacing" gait; that is, they move the two legs on one side of the body before the legs on the other side. This trait is shared with camels and giraffes. As a walk speeds up into a trot, a cat's gait will change to be a "diagonal" gait, similar to other mammals: the diagonally opposite hind and forelegs will move simultaneously.[56]
Like almost all members of the Felidae family, cats have protractable claws.[57] In their normal, relaxed position the claws are sheathed with the skin and fur around the toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hind feet.[58] Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, "kneading", or for extra traction on soft surfaces. Most cats have five claws on their front paws, and four on their rear paws.[59] The fifth front claw (the dewclaw) is proximal to the other claws. More proximally, there is a protrusion which appears to be a sixth "finger". This special feature of the front paws, on the inside of the wrists, is the carpal pad, also found on the paws of big cats and dogs. It has no function in normal walking, but is thought to be an anti-skidding device used while jumping. Some breeds of cats are prone to polydactylyism, and may have eight or even ten toes.[59] These are particularly common along the North-East coast of North America.[60]
Physiology
Normal physiological values[61]
Body temperature
38.6 °C (101.5 °F)
Heart rate
120140 beats per minute
Breathing rate
1640 breaths per minute
As cats are familiar and easily kept animals, their physiology has been particularly well studied; it generally resembles that of other carnivorous mammals but displays several unusual features probably attributable to cats' descent from desert-dwelling species.[26] For instance, cats are able to tolerate quite high temperatures: humans generally start to feel uncomfortable when their skin temperature passes about 44.5 °C (112 °F), but cats show no discomfort until their skin reaches around 52 °C (126 °F),[62] and can tolerate temperatures of up to 56 °C (133 °F) if they have access to water.[63]
Cats conserve heat by reducing the flow of blood to their skin and lose heat by evaporation through their mouth. They do not sweat, and pant only at very high temperatures.[64] Unusually, a cat's body temperature does not vary throughout the day; this is part of cats' general lack of circadian rhythms and may reflect their tendency to be active both during the day and at night.[65] Cats' feces are usually dry and their urine is also highly concentrated, both of which are adaptations that allow cats to retain as much fluid as possible.[26] Their kidneys are so efficient that cats can survive on a diet consisting only of meat, with no additional water,[66] and can even rehydrate by drinking seawater.[67][68]
Cats are obligate carnivores: their physiology has evolved to efficiently process meat, and they have difficulty digesting plant matter.[26] In contrast to omnivores such as rats, which only require about 4% protein in their diet, about 20% of a cat's diet must be protein.[26] Cats are unusually dependent on a constant supply of the amino acid arginine, and a diet lacking arginine causes marked weight loss and can be rapidly fatal.[69] Another unusual feature is that the cat also cannot produce the amino acid taurine, with taurine deficiency causing macular degeneration, where the cat's retina slowly degenerates, causing irreversible blindness.[26] Since cats tend to eat all of their prey, they obtain minerals by digesting animal bones, and a diet composed only of meat may cause calcium deficiency.[26]
A cat's digestive tract is also adapted to meat eating, being much shorter than that of omnivores and having low levels of several of the digestive enzymes that are needed to digest carbohydrates.[70] These traits severely limit the cat's ability to digest and use plant-derived nutrients, as well as certain fatty acids.[70] Despite the cat's meat-oriented physiology, several vegetarian or vegan cat foods have been marketed that are supplemented with chemically synthesized taurine and other nutrients, in attempts to produce a complete diet. However, some of these products still fail to provide all the nutrients that cats require,[71] and diets containing no animal products pose the risk of causing severe nutritional deficiencies.[72]
Senses
Main article: Cat senses
Cats have excellent night vision and can see at only one-sixth the light level required for human vision.[73] This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light.[74] Another adaptation to dim light is the large pupils of cats' eyes. Unlike some big cats, such as tigers, domestic cats have slit pupils.[75] These slit pupils can focus bright light without chromatic aberration, and are needed since the domestic cat's pupils are much larger, relative to their eyes, than the pupils of the big cats.[75] Indeed, at low light levels a cat's pupils will expand to cover most of the exposed surface of its eyes.[76] However, domestic cats have rather poor color vision and (like most non-primate mammals) have only two types of cones, optimized for sensitivity to blue and yellowish green; they have limited ability to distinguish between red and green,[77] although they can achieve this in some conditions.[78]
Cats' whiskers are highly sensitive to touch.
Cats have excellent hearing and can detect an extremely broad range of frequencies. They can hear higher-pitched sounds than either dogs or humans, detecting frequencies from 55 Hz up to 79 kHz, a range of 10.5 octaves; while humans can only hear from 31 Hz up to 18 kHz, and dogs hear from 67 Hz to 44 kHz, which are both ranges of about 9 octaves.[79][80] Cats do not use this ability to hear ultrasound for communication but it is probably important in hunting,[81] since many species of rodents make ultrasonic calls.[82] Cats' hearing is also extremely sensitive and is among the best of any mammal,[79] being most acute in the range of 500 Hz to 32 kHz.[83] This sensitivity is further enhanced by the cat's large movable outer ears (their pinnae), which both amplify sounds and help a cat sense the direction from which a noise is coming.[81]
Cats have an acute sense of smell, which is due in part to their well-developed olfactory bulb and also to a large surface of olfactory mucosa, in cats this mucosa is about 5.8 cm2 in area, which is about twice that of humans and only 1.7-fold less than the average dog.[84] Cats are very sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol,[85] which they use to communicate through urine spraying and marking with scent glands.[86] Cats also respond strongly to plants such as catnip which contains nepetalactone, as they can detect this substance at less than one part per billion.[87] This response is also produced by other plants, such as Silver Vine and valerian, and may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.[88]
Cats have relatively few taste buds compared to humans. Owing to a mutation in an early cat ancestor, one of two genes necessary to taste sweetness may have been lost by the cat family.[38] Their taste buds instead respond to amino acids, bitter tastes and acids.[89] To aid with navigation and sensation, cats have dozens of movable vibrissae (whiskers) over their body, especially their face. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.[90]
Health
Main article: Cat health
In captivity, an average life expectancy for male indoor cats at birth is 12 to 14 years,[91] with females usually living a year or two longer. However, there have been records of cats reaching into their 20s and 30s, with the oldest known cat, Creme Puff, dying at a verified age of 38.[92] Having a cat neutered or spayed confers some health benefits, since castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.[93] The lifespan of feral cats is hard to determine accurately, although one study reported a median age of 4.7 years, with a range between 0 to 8.3 years.[94]
Diseases
Cats can suffer from a wide range of health problems, including infectious diseases, parasites, injuries and chronic disease. Vaccinations are available for many of these diseases, and domestic cats are regularly given treatments to eliminate parasites such as worms and fleas.
Poisoning
In addition to obvious dangers such as rodenticides, insecticides and weed killers, cats may be poisoned by many chemicals that are usually considered safe.[95] This is because their livers are less effective at some forms of detoxification than those of other animals, including humans and dogs.[26][96] Some of the most common causes of poisoning in cats are antifreeze and rodent baits.[97] It has also been suggested that cats may be particularly sensitive to environmental pollutants.[95][98] When a cat has a sudden or prolonged serious illness without any obvious cause, it is possible that it has been exposed to a toxin.
Human medicines should never be given to cats. For example, the painkiller paracetamol (also called acetaminophen, sold as Tylenol and Panadol) is extremely toxic to cats: even very small doses need immediate treatment and can be fatal.[99][100] Even aspirin, which is sometimes used to treat arthritis in cats, is much more toxic to them than to humans and must be administered cautiously.[95] Similarly, application of minoxidil (Rogaine) to the skin of cats, either accidentally or by well-meaning owners attempting to counter loss of fur, has sometimes been fatal.[101] Essential oils can be toxic to cats and there have been reported cases of serious illnesses caused by tea tree oil, and tea tree oil-based flea treatments and shampoos.[102]
Other common household substances that should be used with caution around cats include mothballs and other naphthalene products.[95] Phenol-based products (Pine-Sol, Dettol (Lysol) or hexachlorophene[95]) are often used for cleaning and disinfecting near cats' feeding areas or litter boxes but these can sometimes be fatal.[103] Ethylene glycol, often used as an automotive antifreeze, is particularly appealing to cats, and as little as a teaspoonful can be fatal.[104] Some human foods are toxic to cats; for example theobromine in chocolate can cause theobromine poisoning, although few cats will eat chocolate.[105] Large amounts of onions or garlic are also poisonous to cats.[95] Many houseplants are also dangerous,[106] such as Philodendron species and the leaves of the Easter Lily, which can cause permanent and life-threatening kidney damage.[107]
Behavior
See also: Cat behavior and cat communication
Free-ranging cats are active both day and night, although they tend to be slightly more active at night.[108][109] The timing of cats' activity is quite flexible and varied, which means that house cats may be more active in the morning and evening (crepuscular behavior), as a response to greater human activity at these times.[110] House cats have territories that vary considerably in size, in one study ranging from seven to 28 hectares (69 acres).[109] Although they spend the majority of their time in the vicinity of their home, they can range many hundreds of meters from this central point.[109] Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually 1216 hours, with 1314 being the average. Some cats can sleep as much as 20 hours in a 24-hour period. The term cat nap refers to the cat's ability to fall asleep (lightly) for a brief period and has entered the English lexiconsomeone who nods off for a few minutes is said to be "taking a cat nap". During sleep cats experience short periods of rapid eye movement sleep accompanied by muscle twitches, which suggests that they are dreaming.[111]
Sociability
Social grooming in a pair
Although feral cats are solitary, the social behavior of domestic cats is much more variable and ranges from widely dispersed individuals to feral cat colonies that form around a food source, based on groups of co-operating females.[112][113] Within such groups one cat is usually dominant over the others.[114] Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about ten times larger than those of female cats and may overlap with several females' territories.[86] These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands and by defecation.[86] Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Despite some cats cohabiting in colonies, cats do not have a social survival strategy, or a pack mentality and always hunt alone.[115]
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling, snarling and several different forms of meowing.[8] In contrast, feral cats are generally silent.[116] Their types of body language, including position of ears and tail, relaxation of whole body, and kneading of paws, are all indicators of mood. The tail and ears are particularly important social signal in cats, with a raised tail acting as a friendly greeting.[117][118] Tail raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate animals.[118] Nose-touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.[113] However, some pet cats are poorly socialized. In particular older cats may show aggressiveness towards newly arrived kittens, which may include biting and scratching; this type of behavior is known as Feline Asocial Aggression.[119]
For cats, life in proximity with humans (and other animals kept by humans) amounts to a "symbiotic social adaptation". They may express great affection towards their human companions, especially if they imprint on them at a very young age and are treated with consistent affection. It has been suggested that, ethologically, the human keeper of a cat functions as a sort of surrogate for the cat's mother, and that adult domestic cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny.[120] Conversely, the high-pitched purrs cats make to solicit food may mimic the cries of a hungry human infant, making them particularly hard for humans to ignore.[121]
Grooming
The hooked papillae on a cat's tongue act like a hairbrush to help clean and detangle fur.
Cats are known for their cleanliness, spending many hours licking their coats.[122] The cat's tongue has backwards-facing spines about 500 micrometers long, which are called papillae. These are quite rigid, as they contain keratin.[123] These spines allow cats to groom themselves by licking their fur, with the rows