I was wondering like last week why the price for the 270x wasn't dropping. Normally you'd expect a slow drop in price for this kind of cards. You know, middle range cards...
Did a bit of digging and found out that the 270x is selling really well cuz of their really nice crossfire performance.
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loginR9 270x in crossfire outperforms a R9 290x while - at least for German prices - being cheaper than the 290x.
So, you might go for a R9 270x with the old board and when you wanna upgrade the next time, you'll buy a crossfire board, cpu, psu and another 270x which might have dropped in price till then. I think it's a good upgrade strategy. The gpu is confirmed to be partially DX12 compatible too, not for every single shader maybe but for the "big" improvement due to that "mantle" thing they wanna deliver (I know it's called differently but I am too lazy to google).
I am using a XFX R9 270x with 2GB atm. Witcher 3 on full Ultra except Hairworks and ViewDistance, Vsync on: steady 30 fps without any hickup...
It's a solid card for the "small" money. I'd go for the 4GB variant tho. Some games already use that much (GTA 5)